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Description
They are counterparts of the Knuckles that appear throughout history.

O'Nux: also known as Anti Knuckles is a Moebian Echidna and Knuckles's alternate self from Moebius. Just as his counterpart is Guardian of Angel Island, O'Nux is the Overseer of Demon Island. But most unlike Knuckles, he is a pacifist who only uses violence as a last resort. (StH: #44). He was defeated when Scourge the Hedgehog took over Moebius with the aid of the Suppression Squad. (StH: #44)
Sir Gawain: is a Mobian echidna and the alternate counterpart of Knuckles the Echidna from World of Camelot, as Gawain and the Knights of the Round Table saw King Arthur lose Sonic the Hedgehog and Merlina, the trio of knights were ordered by Arthur to search for Merlina and Sonic and slay them. After Arthur left, Gawain voiced his objections to their orders, as Merlina was the royal wizard. Lancelot and Percial, however, reminded him of the importance of obeying their king, and Gawain went on his mission (StH: #197)
Knuckles the Echidna (no Knuckle spikes): One of the prisoners in the zone jail within the No Zone, it is unknown about his Zone of origin and if he also shares the same name as his alternate version of Mobius, he was probably one of the prisoners who liked to provoke Scourge, because he used Scourge as a basketball (SU: #29), later on Scourge's escape, Scourge finds him and says if he is the echidna that made him a basketball that day, even denying it, Scourge knocked him out with a Spin-Dash on the belly (SU: #32).
Knuckles (Crystal Claws): Not much is known about him, nor about his Zone of origin and if he also shares the same name as his alternate version of Mobius, he appears with the other prisoners in a therapy section with Zouge, he soon afterwards agrees with the other prisoners who all deserve revenge (SU #31). During the riot, he was seen attempting to escape. (SU #32).
Knuckson: is an alternate counterpart of Knuckles the Echidna from the Sentai Zone, Knuckson and the rest of the Sonicaman: Chaos Ninja Team were summoned to Mobius Prime by Zonic the Zone Cop to save Sonic from Sallactor's attacks. (SSS: #12)
Lieutenant Knuckles: Was an alternate version of Knuckles the Echidna from the Litigopolis Zone, he was one of the many that had mistaken Sonic for Anti Sonic. When he was about to arrest Sonic, he was knocked out by Sally McAcorn. (SSS: #14)
Knuckles-Zilla: Is an alternate version of Knuckles the Echidna from an alternate version of Mobius. In this universe, Chaos Emerald radiation transformed Knuckles into a hideous, gigantic monster that threatened the people of Mayor Kintobor's city. Despite his fearsome mutatations, he was unable to stop the Giant Robotno piloted by Sonic the Hedgehog. (SSS: #12)
Tuxedo Knux: He was the alternate counterpart to Knuckles the Echidna from the Luna Zone, after Ivanna Robotina escaped their zone and briefly attacked Mobius Prime, Tuxedo Knux pursued her along with Sally Moon and Chibi Rose. Following a brief scuffle he rendered Robotina unconscious using one of his roses, and returned her to his native zone with the others, just after sharing a romantic moment with Sally Moon. (SSS: #8).

Sir Gawain is from SEGA, the other characters are from Archie Comics

Credits:
nagoyamonkey-smg by Tuxedo Mask Sprites
SonicKnight98 Sir Gawain Sprites
earthbouds

Description
They are counterparts of the Knuckles that appear throughout history.

O'Nux: also known as Anti Knuckles is a Moebian Echidna and Knuckles's alternate self from Moebius. Just as his counterpart is Guardian of Angel Island, O'Nux is the Overseer of Demon Island. But most unlike Knuckles, he is a pacifist who only uses violence as a last resort. (StH: #44). He was defeated when Scourge the Hedgehog took over Moebius with the aid of the Suppression Squad. (StH: #44)
Sir Gawain: is a Mobian echidna and the alternate counterpart of Knuckles the Echidna from World of Camelot, as Gawain and the Knights of the Round Table saw King Arthur lose Sonic the Hedgehog and Merlina, the trio of knights were ordered by Arthur to search for Merlina and Sonic and slay them. After Arthur left, Gawain voiced his objections to their orders, as Merlina was the royal wizard. Lancelot and Percial, however, reminded him of the importance of obeying their king, and Gawain went on his mission (StH: #197)
Knuckles the Echidna (no Knuckle spikes): One of the prisoners in the zone jail within the No Zone, it is unknown about his Zone of origin and if he also shares the same name as his alternate version of Mobius, he was probably one of the prisoners who liked to provoke Scourge, because he used Scourge as a basketball (SU: #29), later on Scourge's escape, Scourge finds him and says if he is the echidna that made him a basketball that day, even denying it, Scourge knocked him out with a Spin-Dash on the belly (SU: #32).
Knuckles (Crystal Claws): Not much is known about him, nor about his Zone of origin and if he also shares the same name as his alternate version of Mobius, he appears with the other prisoners in a therapy section with Zouge, he soon afterwards agrees with the other prisoners who all deserve revenge (SU #31). During the riot, he was seen attempting to escape. (SU #32).
Knuckson: is an alternate counterpart of Knuckles the Echidna from the Sentai Zone, Knuckson and the rest of the Sonicaman: Chaos Ninja Team were summoned to Mobius Prime by Zonic the Zone Cop to save Sonic from Sallactor's attacks. (SSS: #12)
Lieutenant Knuckles: Was an alternate version of Knuckles the Echidna from the Litigopolis Zone, he was one of the many that had mistaken Sonic for Anti Sonic. When he was about to arrest Sonic, he was knocked out by Sally McAcorn. (SSS: #14)
Knuckles-Zilla: Is an alternate version of Knuckles the Echidna from an alternate version of Mobius. In this universe, Chaos Emerald radiation transformed Knuckles into a hideous, gigantic monster that threatened the people of Mayor Kintobor's city. Despite his fearsome mutatations, he was unable to stop the Giant Robotno piloted by Sonic the Hedgehog. (SSS: #12)
Tuxedo Knux: He was the alternate counterpart to Knuckles the Echidna from the Luna Zone, after Ivanna Robotina escaped their zone and briefly attacked Mobius Prime, Tuxedo Knux pursued her along with Sally Moon and Chibi Rose. Following a brief scuffle he rendered Robotina unconscious using one of his roses, and returned her to his native zone with the others, just after sharing a romantic moment with Sally Moon. (SSS: #8).

Sir Gawain is from SEGA, the other characters are from Archie Comics

Credits:
nagoyamonkey-smg by Tuxedo Mask Sprites
SonicKnight98 Sir Gawain Sprites
earthbouds